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Sixth stage of the Permian
timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between
Wordian
Second series and epoch of the Permian
more information on this topic. The Wordian Stage was between 268.8 ± 0.5 – 265.1 ± 0.4 Mya. The base of the Wordian Stage is defined as the place in the
Guadalupian
Sixth and last period of the Paleozoic Era
Lopingian, Capitanian and part of the Wordian, while the underlying Kazanian includes the rest of the Wordian as well as the Roadian.[needs update] In
Permian
Fifth stage of the Permian
the Kungurian and followed by the Wordian. In 1961, the regional timescale used for the southeastern US had the Wordian and Capitanian as subdivisions of
Roadian
Geological formation in Brazil
publications it appears as Rio do Rastro. The strata were deposited between the Wordian and the Wuchiapingian, from about 266 to 254 million years ago. The geology
Rio_do_Rasto_Formation
Seventh stage of the Permian geologic period
lasted between 264.28 and 259.857 million years ago. It was preceded by the Wordian and followed by the Wuchiapingian. A significant mass extinction event
Capitanian
Extinct group of saber-toothed therapsids from the Permian
fossil record, constrained to a minimum age before the middle Wordian, if not pre-Wordian. This suggests that the absence of gorgonopsians from low-latitudes
Gorgonopsia
Family of annelids
live in saline waters. The earliest serpulids are known from the Permian (Wordian to late Permian). The blood of most species of serpulid and sabellid worms
Serpulidae
Extinct genus of cartilaginous fishes
hybodontiform. It lived from the Tournaisian age of the Early Carboniferous to the Wordian age of the Permian, and remains have been found in England, Russia and
Reesodus
Extinct genus of therapsid reptile
Ivantosaurus is an extinct genus of therapsid that lived in Russia during the Wordian stage of the Permian period, named in honor of paleontologist Ivan Antonovich
Ivantosaurus
Genus of molluscs (fossil)
1887: Wordian of Italy Prostacheoceras burnense Tumanskaia, 1931: Roadian of Russia (Crimea) Prostacheoceras chinense Wang, 1983: Asselian - Wordian of China
Prostacheoceras
Extinct family of vertebrate animals
from the strata of the middle Permian (upper Roadian to lower Wordian) and (latest Wordian-earliest Capitanian) of France, where they cohabited notably
Caseidae
Extinct family of therapsids
widely distributed across Pangaea. They flourished briefly during the Wordian and Capitanian ages, radiating into several lineages, existing simultaneously
Tapinocephalidae
Extinct genus of synapsids
caseid synapsid that lived during the Middle Permian (late Roadian - early Wordian) in northern European Russia. The genus is only represented by its type
Ennatosaurus
Dzhulfa Formation Julfa Formation Khachik Formation Capitanian Arpinskaya Formation Wordian-Capitanian Gnishik Formation Wordian Arpa Formation Wordian
List of fossiliferous stratigraphic units in Armenia
List_of_fossiliferous_stratigraphic_units_in_Armenia
Extinct genus of therapsid
gorgonopsian therapsid (mammal-line synapsid) known from the Permian (Wordian–Capitanian age) Abrahamskraal Formation (Tapinocephalus Assemblage Zone)
Jirahgorgon
Extinct family of therapsids
therapsids. The Eotitanosuchidae were large predatory therapsids of the Wordian epoch. It was once considered to belong to a separate infraorder of therapsids
Eotitanosuchidae
Russia. Specimens of Amelacanthus have additionally been found in Permian (Wordian) and Triassic (Induan) sediments of Oman. This makes Amelacanthus one of
Amelacanthus
Group of mites
Karoo locality within the Karoo Supergroup in South Africa, dating to the Wordian stage of the Middle Permian, approximately 266 million years ago. Di Sabatino
Hydrachnidia
Extinction event around 260 million years ago
event but a continuous decline in diversity that began at the end of the Wordian. Another study examining fossiliferous facies in Svalbard found no evidence
Capitanian mass extinction event
Capitanian_mass_extinction_event
Order of insects
Russia, Middle Permian (Wordian) †Zoropsocus Tillyard, 1935 Kazankovo-Markinskaya Formation, Russia, Middle Permian (Wordian) Croudace Bay Formation,
Lophioneurida
Extinct genus of amphibians
Sterlitamak site in Bashkortostan in Russia. The span of the strata is listed as Wordian (268–265.8 million years ago). Melosaurus was 2.5–3 m (8.2–9.8 ft) in length
Melosaurus
System that relates geologic strata to time
Wuchiapingian 259.857 ± 0.21 * Guadalupian Capitanian 264.28 ± 0.16 * Wordian 266.9 ± 0.4 * Roadian 274.4 ± 0.4 * Cisuralian Kungurian 283.3 ± 0.4 Artinskian
Geologic_time_scale
Ninth and last stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Changhsingian
Extinct genus of ammonites
Elephantoceras Temporal range: Wordian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Mollusca Class: Cephalopoda Subclass:
Elephantoceras
Geological group in South Africa
the Beaufort Group rocks are considered to range between Middle Permian (Wordian) to Early Triassic (Anisian) in age. During the period when sedimentation
Beaufort_Group
Dzhulfa Formation Julfa Formation Khachik Formation Capitanian Arpinskaya Formation Wordian-Capitanian Gnishik Formation Wordian Arpa Formation Wordian
List of fossiliferous stratigraphic units in Azerbaijan
List_of_fossiliferous_stratigraphic_units_in_Azerbaijan
Subclass of fishes
Twitchett, Richard J. (March 2013). "A new diverse shark fauna from the Wordian (Middle Permian) Khuff Formation in the interior Haushi-Huqf area, Sultanate
Elasmobranchii
County in Texas, United States
for rock strata in Road Canyon in the northern part of the county. The Wordian Age of the Permian Period is probably named for the Word Ranch in the Glass
Brewster_County,_Texas
Extinct genus of therapsids
mother, Pamela. Fossils have been found dating from the Middle Permian (Wordian age). Tapinocaninus fossils were first found in the Eodicynodon Assemblage
Tapinocaninus
Extinct superfamily of reptiles
the middle Pennsylvanian to the middle Guadalupian epoch (Moscovian - Wordian stages) of Europe, North America and Asia. It was named by the Russian
Lanthanosuchoidea
Extinct genus of bryozoans
H. (November 2014). "Bryozoa of the uppermost Gerster Limestone (late Wordian, Permian), Medicine Range, northeastern Nevada, U.S.A.". Rocky Mountain
Utgaardostylus
Extinct family of beetles
Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District
Permocupedidae
Extinct genus of synapsids
the Middle Wordian 266.66 ± 0.76 million years ago. Therefore, this part of the formation would probably have a late Roadian – early Wordian age, while
Lalieudorhynchus
Extinct clade of insects
Gonfaron, Pélitique Formation, France, Early–Middle Permian (Artinskian–Wordian) (known from a wing fragment) incertae sedis †Ditaxineurella Martynov,
Protanisoptera
Extinct genus of therapsids
Pachydectes was present solely during the Guadalupian, specifically during the Wordian and Capitanian without being found anywhere else. Pachydectes is a genus
Pachydectes
Eighth stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Wuchiapingian
Extinct clade of synapsids
successful therapsids. The last caseasaurs became extinct between the latest Wordian and earliest Capitanian stage of the Middle Permian. Among the most conspicuous
Caseasauria
First series of the Permian
Guadalup Lopin Early Triassic Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian ← Permian-Triassic mass extinction
Cisuralian
Extinct genus of therapsids
family Anteosauridae. It lived 268 to 265 million years ago during the Wordian age of the Guadalupian (= middle Permian) period in what is now Brazil
Pampaphoneus
Extinct genus of jawless fishes
the species Jinogondolella nanginkensis first appeared. The base of the Wordian stage is defined as the place in the stratigraphic record where fossils
Jinogondolella
Extinct genus of fishes
following species are known: †A. barbarus Minikh, 2006 - Middle Permian (Wordian) of Arkhangelsk, Russia †A. chuvashovi Yankevich, 2001 - Early Permian
Acrolepis
Geologic group in Europe
Guadalupian Lopingian Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Zechstein Rotliegend Cisuralian
Rotliegend
First stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Asselian
Extinct genus of therapsids
Zone was part of the Dinocephalian tetrapod fauna, dating from middle the Wordian to the middle Capitanian stage of the Middle Permian. The assemblage zone
Titanophoneus
Extinct genus of bryozoans
H. (November 2014). "Bryozoa of the uppermost Gerster Limestone (late Wordian, Permian), Medicine Range, northeastern Nevada, U.S.A.". Rocky Mountain
Wyseotrypa
Order of annelids
Sabellida Temporal range: Middle Permian (Wordian)-Present PreꞒ Ꞓ O S D C P T J K Pg N Possible Miaolingian record Bispira sp. Scientific classification
Sabellida
Extinct family of molluscs
Mississippian–Middle Permian PreꞒ Ꞓ O S D C P T J K Pg N Daraelites, a Wordian daraelitid Scientific classification Kingdom: Animalia Phylum: Mollusca
Daraelitidae
Third stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Artinskian
Extinct genus of synapsids
Ennatosaurus tecton, Arkhangelsk Oblast, Russia, late Roadian – early Wordian; 3 Phreatophasma aenigmaticum, Bashkortostan, Russia, early Roadian; 4
Angelosaurus
Extinct family of therapsids
from the fossil record that extends at least as far back as them into the Wordian stage of the Middle Permian, roughly between 266.9 and 264.7 million years
Lycosuchidae
Group of extinct animals
Formation of Mallorca in the western Mediterranean dating to earliest Wordian of the Middle Permian, although the stratigraphic position of the gorgonopsian
Saber-toothed_predator
Extinct genus of synapsids
Formation and the Octon Member of the Salagou Formation) and the late Roadian-Wordian and possibly early Capitanian (for La Lieude Formation). A more precise
Euromycter
Third and final series of the Permian
Guadalup Lopin Early Triassic Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian ← Permian-Triassic mass extinction
Lopingian
Moreno-Bedmar, J. A.; Utrup, J. (2025). "Permian ammonoids from the Guadalupian (Wordian–Capitanian) of Las Delicias, Coahuila state, Mexico: new biostratigraphical
2025_in_paleomalacology
Fourth stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Kungurian
Extinct superfamily of ammonites
Thalassoceratoidea Temporal range: Bashkirian to Wordian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Mollusca Class:
Thalassoceratoidea
Permocallipteris Gen. et comb. nov Valid Naugolnykh Permian (Artinskian to Wordian) China Mongolia Russia A seed fern belonging to the family Peltaspermaceae
2014_in_paleobotany
Genus of conodonts
species M. daheshenensis and M. subgracilis are from the Permian (probably Wordian) of the Daheshen Formation in China. The top of the Sakmarian stage (the
Mesogondolella
Extinct genus of synapsids
Formation and the Octon Member of the Salagou Formation) and the late Roadian-Wordian and possibly early Capitanian (for La Lieude Formation). A more precise
Ruthenosaurus
Extinct genus of dicynodonts
roughly constrained to between 266.9 and 264.7 million years old during the Wordian stage of the middle Permian period (Guadalupian). NMQR 2913 consists of
Nyaphulia
Extinct genus of therapsids
a Middle Permian (or Guadalupian) age, dating to somewhere within the Wordian to Capitanian stages. At the time of its description, Phorcys represented
Phorcys_(therapsid)
South Korean computer program
Japanese (1996) Hangul 97, 97 strengthen, 815 special edition (1998) Hangul Wordian, Hangul for Kids (2000) Hangul 2002 (2001, widely used for government e-document
Hangul_(word_processor)
Ecosystem
for the Old Word Ranch in the Glass Mountains of Brewster County. The Wordian age of the Permian Period is named for the Word Formation. The formation
Word_Formation
Extinct genus of synapsids
younger rocks (ichnogenus Merifontichnus from the La Lieude Formation (Wordian) in the Lodève basin). Remains of Alierasaurus were discovered grouped
Alierasaurus
Extinct subfamily of therapsids
S D C P T J K Pg N It is possible that they developed as early as the Wordian era. Visualized head of Inostrancevia latifrons. Scientific classification
Inostranceviinae
Extinct genus of therapsids
yet therapsids did not appear as a diverse group until near the Roadian-Wordian boundary. This has left a morphological and temporal gap in the fossil
Raranimus
younger than other similar Lagerstätten within the region. Word Formation Wordian Texas, US A proposed "Liberation Lagerstätte".see notes Fossils recovered
List_of_lagerstätten
Use of a particular fossil class to identify stratigraphy
Roadian (the base of the Wordian stage) is at the first appearance of fossils of Jinogondolella aserrata. The base of the Wordian stage is defined as the
Conodont_biostratigraphy
Geologic formation in Mallorca, Spain
overlying Pedra de s'Ase Formation was dated to the middle Permian (Roadian–Wordian) by a particular palynological association and paleomagnetic data. A year
Port_des_Canonge_Formation
Extinct genus of therapsids
Biarmosuchus Temporal range: Wordian, 267 Ma PreꞒ Ꞓ O S D C P T J K Pg N Mounted skeleton of B. tener (above) and restoration (below) Scientific classification
Biarmosuchus
Extinct genus of reptiles
"forgotten butcher") is an extinct genus of early reptile from Middle Permian (Wordian stage, or uppermost Kazanian in Eastern Europe) deposits of Arkhangelsk
Lanthanolania
Second stage of the Permian
Guadalup Lopin ET Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Induan ← Permian-Triassic
Sakmarian
Extinct genus of therapsids
Estemmenosuchus Temporal range: Guadalupian (Wordian), 267 Ma PreꞒ Ꞓ O S D C P T J K Pg N E. uralensis skeleton Scientific classification Kingdom: Animalia
Estemmenosuchus
Group Formation Period Notes Maputo Formation Late Aptian Lupata Group Aptian Lebombo Formation Middle Jurassic Lower Karoo Formation Wordian
List of fossiliferous stratigraphic units in Mozambique
List_of_fossiliferous_stratigraphic_units_in_Mozambique
Extinct genus of moss animals
bryozoans in the family Nematotrypidae. The species C. elegans is from a Wordian (Permian) marine horizon in the Sijiashan Formation of Northeast China
Clausotrypa
Genus of molluscs (fossil)
1937), formerly placed into genus Paraceltites. Its fossils were found in Wordian (Permian) of Texas. "Goniat database". Retrieved 23 September 2015. T.
Anatsabites
Extinct order of chondrichthyans
Tintori, A., and Twitchett, R. J., 2013, A new diverse shark fauna from the Wordian (Middle Permian) Khuff Formation in the interior Haushi-Huqf area, Sultanate
Hybodontiformes
Geologic formation in Namibia
The Gai-As Formation is an Early to Middle Permian (Kungurian to Wordian) geologic formation correlated with the Ecca Group and designated "Ecca" Group
Gai-As_Formation
Omingonde Formation Anisian-Ladinian "Ecca" Group Gai-As Formation Kungurian-Wordian Huab Formation Artinskian-Kungurian Ecca Group Whitehill Formation Artinskian-Kungurian
List of fossiliferous stratigraphic units in Namibia
List_of_fossiliferous_stratigraphic_units_in_Namibia
Geologic formation in New Mexico and Texas, US
Cherry Canyon Formation Stratigraphic range: Roadian-Wordian ~272–266 Ma PreꞒ Ꞓ O S D C P T J K Pg N Cherry Canyon Formation exposed in roadcut near its
Cherry_Canyon_Formation
Genus of therapsid from the Middle Permian of South Africa
Tapinocephalus (Capitanian age, 265.8-260.4 Ma), and the base of the Eodicynodon (Wordian age, 268–265.8 Ma) assembly zones, of the Karoo Basin. Older classifications
Ictidosaurus
Extinct genus of sponges
Guadalupia is an extinct genus of sea sponges. It includes a number of extinct species including: Guadalupia auricula, G. cupulosa, G. ramescens, G. microcamera
Guadalupia
Permian geologic formation in Transcaucasia
formation in Armenia, Azerbaijan and Iran. It preserves fossils dated to the Wordian age of the Permian period. The thin-bedded limestones of the formation
Gnishik_Formation
fauna from the Wordian (Middle Permian) Khuff Formation in the interior Haushi-Huqf area, Sultanate of Oman: CHONDRICHTHYANS FROM THE WORDIAN KHUFF FORMATION
1985_in_paleontology
Extinct family of insects
Russia, Roadian; Kazankovo-Markinskaya Formation, Russia, Middle Permian (Wordian); Mal'tseva Formation, Russia, Changhsingian †Sukhonia Aristov 2013 – Poldarsa
Geinitziidae
Extinct genus of therapsids
lower Permian, dominated by the pelycosaurs, and the mid-middle Permian (Wordian), where the therapids take the first place in terrestrial ecosystems. The
Sinophoneus
Extinct clade of therapsids
Dicynodontia Temporal range: Middle Permian (Wordian) to Late Triassic (late Norian or Rhaetian) 268–211 Ma PreꞒ Ꞓ O S D C P T J K Pg N Cisuralian Guad
Dicynodontia
Permian rock layers between the east coast of England and northern Poland
Guadalupian Lopingian Gzhelian Asselian Sakmarian Artinskian Kungurian Roadian Wordian Capitanian Wuchiapingian Changhsingian Zechstein Rotliegend Cisuralian
Zechstein
Permian geologic formation in Transcaucasia
formation in Armenia and Azerbaijan. It preserves fossils dated to the Wordian to Capitanian ages of the Permian period. List of fossiliferous stratigraphic
Arpinskaya_Formation
Extinct subfamily of therapsids
Anteosaurinae Temporal range: Middle Permian (Wordian to Capitanian), ~266–260 Ma PreꞒ Ꞓ O S D C P T J K Pg N As. S Artin. Kung. Road. W Ca. Wu. C Skeleton
Anteosaurinae
Geologic formation in the Middle East
fauna from the Wordian (Middle Permian) Khuff Formation in the interior Haushi-Huqf area, Sultanate of Oman: CHONDRICHTHYANS FROM THE WORDIAN KHUFF FORMATION
Khuff_Formation
Sphenacanthus riorastoensis Sp. nov Valid Pauliv, Dias & Sedor Permian (Wordian to Wuchiapingian) Rio do Rasto Formation Brazil A species of Sphenacanthus
2012_in_paleoichthyology
Extinct family of beetles
species) – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) Mitina Formation, Kemerovo, Russia, Middle Permian (Roadian) Tecticupes
Taldycupedidae
Extinct genus of therapsids
specimen coming from the Mediterranean island of Mallorca, dating to at least Wordian stage of the Middle Permian, with the possibility of dating to the Early
Rubidgea
Sakmarian Mount Glossopteris Formation Lopingian Bainmedart Coal Measures Wordian Polarstar Formation Guadalupian Radock Formation Kungurian Weller Coal
List of fossiliferous stratigraphic units in Antarctica
List_of_fossiliferous_stratigraphic_units_in_Antarctica
paucinervis. Pycnoblattina valeriea Sp. nov Garrouste, Boderau & Nel Permian (Wordian) France A roachoid belonging to the family Spiloblattinidae. Recyklovany
2025_in_paleoentomology
Extinct superfamily of ammonites
Tournaisian – Bashkirian? stages) and the derived Daraelitidae (Viséan – Wordian stages). Prolecanitids and daraelitids differ primarily in the complexity
Prolecanitoidea
Geologic formation in Oregon, US
Cisuralian Series Sakmarian formations Artinskian formations Roadian formations Wordian formations Kungurian formations Permian North America Esplanade Sandstone
Coyote_Butte_Limestone
Genus of anteosaurid synapsid from the Permian
biostratigraphically subdivided in two faunal zone: the older, mostly Wordian-age Eodicynodon Assemblage Zone, and the younger Tapinocephalus Assemblage
Anteosaurus
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