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Geologic formation in Utah notable for trilobite fossils
Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale,
Wheeler_Shale
Fine-grained, clastic sedimentary rock
Places Oil shale – Organic-rich fine-grained sedimentary rock containing kerogen Shale gas – Natural gas trapped in shale formations Wheeler Shale – Geologic
Shale
"Pseudoarctolepis sharpi, n. gen., n. sp. (Phyllocarida), from the Wheeler Shale (Middle Cambrian) of Utah". Journal of Paleontology. 30 (1): 9–14. ISSN 0022-3360
List_of_Cambrian_arthropods
Fossil-bearing rock formation in the Canadian Rockies
Maotianshan Shales, which is often compared to Burgess Shale Paleobiota of the Burgess Shale Wheeler Shale, also compared to Burgess Shale List of arthropods
Burgess_Shale
Class of extinct, Paleozoic arthropods
well-known Elrathia kingi trilobite is found in abundance in the Cambrian Wheeler Shale of Utah. Spectacularly preserved trilobite fossils, often showing soft
Trilobite
Genus of Cambrian animals
lobopodian known from Cambrian-aged fossils in Burgess Shale-type deposits in Canada (Burgess Shale) and China, and from isolated spines around the world
Hallucigenia
Extinct genus of Cambrian Era Chordate
Sirius Passet, Banffia of the Burgess Shale, and Skeemella of the Pierson Cove Formation above the Wheeler Shale). N. aldridgei is known from several incomplete
Nesonektris
Genus of trilobites
fossils in the USA locally found in extremely high concentrations within the Wheeler Formation in the U.S. state of Utah. E. kingii has been considered the
Elrathia
Extinct genus of soft-bodied animals
prolata was described as a separate species from the similarly-aged Wheeler Shale Formation of Utah but in fact represents the frontal appendage of a
Aysheaia
Extinct genus of Cambrian animals
known from the Middle Cambrian Burgess Shale, the Comley limestone, the Wheeler Shale, the Bright Angel Shale and elsewhere (such as Iran). It is named
Chancelloria
Extinct genus of arthropods
species T. palaeothalassia is known from the Burgess Shale, Canada T. holmani is from the Wheeler Shale of Utah, Thelxiope tangi from the Linyi Lagerstätte
Thelxiope
Extinct genus of arthropod
Cambropodus is a genus of Cambrian arthropod from the Wheeler Shale of Utah. It contains one species, Cambropodus gracilis. Cambropodus is only known
Cambropodus
Extinct genus of marine invertebrates
Notable localities where species are found include the Wheeler Shale of Utah, and the Burgess Shale of British Columbia.[citation needed] The species of
Gogia
Extinct genus of Cambrian radiodont
(particularly the Burgess Shale) of British Columbia, Canada. The other named species A. daleyae is known from the somewhat older Emu Bay Shale of Australia. Other
Anomalocaris
Extinct genus of arthropods
species was done in 2012. A second species E. brutoni is known from the Wheeler Shale, which was described in 2011. An additional specimen of E. brutoni was
Emeraldella
Extinct genus of bivalved arthropod
primarily the Burgess Shale of British Columbia, Canada but also possibly the Wheeler Shale, Marjum Formation, Pioche Shale and Bloomington Formation
Perspicaris
Genus of sponges
Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; the Itajaí Basin in Brazil;
Choia
Genus of crustaceans
best known species, Branchiocaris pretiosa, was described from the Burgess Shale of British Columbia, Canada, in 1929, originally placed in Protocaris, and
Branchiocaris
Extinct species of trilobite
as an index fossil. Its first appearance at the GSSP section in the Wheeler Shale of Utah is defined as the beginning of the Drumian Age (around 504.5
Ptychagnostus_atavus
Extinct genus of priapulid worms
spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited 508 million years ago. Microfossils
Ottoia
Genus of vetulocystids
Thylacocercus is a monotypic genus of vetulocystid from the Wheeler Shale of Utah that is tentatively placed within the family Vetulocystidae. Its discovery
Thylacocercus
Extinct genus of arthropods
belonging to C. perfecta, are also found in the Spence Shale of western Utah as well as the Pioche Shale of Nevada. Canadaspis laevigata comes from the Chengjiang
Canadaspis
Extinct genus of bilaterians
Skeemella is a genus of elongate animal from the Middle Cambrian Wheeler Shale and Marjum lagerstätte of Utah. It has been classified with the banffozoan
Skeemella
Extinct genus of arthropods
Early and Middle Cambrian. Known from over 300 specimens in the Burgess Shale and the Chengjiang biota. It is a member of the family Leanchoiliidae in
Alalcomenaeus
Extinct genus of arthropods
Dicranocaris is an extinct genus of arthropod that lived in the Wheeler Shale formation of what is now modern day Utah during Drumian of the Cambrian
Dicranocaris
Genus of Cambrian animals
corrugata, which is known from hundreds of complete specimens in the Burgess Shale; other species are known only from fragmentary material or limited sample
Wiwaxia
Extinct genus of arthropods
Columbia, Canada. When the fossil was first discovered in Canada's Burgess Shale, it was believed to be a crustacean, such was the difference between this
Naraoia
Genus of trilobites
soft body parts intact in the Burgess Shale in Canada. Fossils of this genus can also be found in the Wheeler Shale and Marjum Formation of Utah in the
Olenoides
Extinct genus of sponges
Crumillospongia is a genus of middle Cambrian sponges known from the Burgess Shale and other localities from the Lower and Middle Cambrian. Its name is derived
Crumillospongia
Geological formation in Utah, United States
Marjum Formation is a Cambrian geological formation that overlies the Wheeler Shale in the House Range, Utah. It is named after its type locality, Marjum
Marjum_Formation
Extinct genus of trilobites
Hillard Peak, Alaska; lower part of the Bolaspidella-zone, Western-Utah; Wheeler Shale, Antelope Spring, House Range, Utah), the Russian Federation (Malokuonamsky
Itagnostus
Extinct genus of arthropod
Canada, Parker Shale, Vermont, Pioche Shale, Nevada Kinzers Formation, Pennsylvania, USA Tuzoia australis Glaessner, 1979 Emu Bay Shale, Australia Tuzoia
Tuzoia
Extinct genus of radiodonts
It is known by a single species, Pahvantia hastata, described from Wheeler Shale and Marjum Formation in Utah. Although it was once considered as a filter
Pahvantia
Extinct genus of arthropods
megacheiran marine arthropod known from Cambrian deposits of the Burgess Shale in Canada and the Chengjiang biota of China. L. superlata was about 5 centimetres
Leanchoilia
Genus of extinct arthropods
Jago, Gehling & Lee, 2009 Emu Bay Shale, Australia, Cambrian Stage 4 Isoxys communis Glaessner, 1979 Emu Bay Shale, Australia, Cambrian Stage 4 Isoxys
Isoxys
Genus of extinct arthropods
from the Early to the Mid Cambrian of China and the Mid Cambrian Burgess Shale of British Columbia, Canada. It is thought to have been a durophagous (crushing)
Sidneyia
Cambrian fossil genus
Margaretia is a frondose organism known from the middle Cambrian Burgess Shale and the Kinzers Formation of Pennsylvania. Its fronds reached about 10 cm
Margaretia
Extinct genus of stem-chelicerates
not described as a new genus until 2026. The fossil originates from the Wheeler Formation (Miaolingian: Drumian) of Utah. The genus name is intended to
Megachelicerax
Genus of marine worm-like animals
Yuknessia is an early pterobranch, known from the Burgess Shale and the Wheeler Shale. Long, unbranched fronds emerge from a central holdfast-like body
Yuknessia
Cambrian of Australia, Canada (British Columbia, especially in the Burgess Shale, and Newfoundland), Greenland, Mexico, and the United States (Alaska, Idaho
Bathyuriscus
Extinct order of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Proetida
Cambrian arthropod
known from the Burgess Shale Lagerstätte of British Columbia, Canada. Specimens of Waptia are also known from the Spence Shale of Utah, United States
Waptia
Extinct genus of trilobites
Olenoides has. Bathyuriscus (Kootenia) dawsoni Walcott, 1889. From the Burgess shale (the Stephen formation) of British Columbia. Kootenia is sometimes believed
Kootenia
Extinct genus of soft-bodied animals
soft-bodied cambroernid best known from the Fossil Ridge outcrops of the Burgess Shale, particularly in the 'Great Eldonia layer' in the Walcott Quarry. In addition
Eldonia
Fossilized fauna of Burgess shale
to that of the Burgess Shale. While many are also preserved in a similar fashion to the Burgess Shale, the term "Burgess Shale-type fauna" covers assemblages
Burgess_Shale-type_fauna
Extinct genus of bivalved arthropod
was described by Brooks & Caster in 1956 from specimens found in the Wheeler Shale of Utah. It is unusual among Cambrian arthropods for having a pair of
Pseudoarctolepis
have been found in Canada (British Columbia, especially in the Burgess Shale, and Newfoundland), Greenland, Mexico, and the United States (Alaska, Nevada
Zacanthoides
Extinct genus of gilled lobopodian
found in Utah, although an origin in the Weeks Formation or most of the Wheeler Shale can be ruled out due to differing fossil preservation methods. McCall
Utahnax
Genus of trilobites
the United States (Delamaran, Lower Wheeler Shale, Millard County, Utah, 40.0°N, 113.0°W; and Menevian, Wheeler Formation, House Range, Utah, 39.2° N
Asaphiscus
Extinct genus of trilobites
punctuosus Angelin, 1851 from the Ptychagnostus punctuosus zone of the Alum Shale (Drumian), Sweden (by original designation). Official ruling on the conservation
Ptychagnostus
Fossil flora and fauna of the Burgess Shale
list of the biota of the Burgess Shale, a Cambrian lagerstätte located in Yoho National Park in Canada. The Burgess Shale is a fossil-bearing deposit exposed
Paleobiota of the Burgess Shale
Paleobiota_of_the_Burgess_Shale
Extinct genus of sponges
Diagoniella is a genus of sponge known from the Middle Cambrian Burgess Shale. 128 specimens of Diagoniella are known from the Greater Phyllopod bed,
Diagoniella
Mountain range in Millard County, Utah, USA
range is known for a fossil Lagerstätte (Wheeler Shale) of Cambrian age, which has an array of Burgess Shale type fauna, including Elrathia kingii, a
House_Range
Extinct genus of marine invertebrates
Cothurnocystis is a genus of small enigmatic echinoderms that lived during the Ordovician. Individual animals had a flat boot-shaped body and a thin rod-shaped
Cothurnocystis
Extinct genus of priapulid worms
tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale, Puncoviscana Formation, the Early Ordovician Fezouata Formation
Selkirkia
American baseball player (born 1990)
Kennesaw State. Wheeler played for the Giants' minor league teams in 2010 and 2011 before he was traded to the Mets in July 2011. Wheeler then rose through
Zack_Wheeler
Extinct genus of bacteria
as carbonaceous films in the Middle Cambrian Burgess Shale. it is present throughout the shale; 2580 specimens of Morania are known from the Greater
Morania
Extinct order of arthropods
2019 study of adult specimens with preserved soft tissue from the Burgess Shale found that agnostidans shared morphological similarities to trilobites and
Agnostida
Extinct genus of trilobites
Cedaria is an extinct genus of trilobites from the late Cambrian. It is a small, rather flat trilobite with an oval outline, a headshield and tailshield
Cedaria
Extinct genus of jawless fishes
conodonts in the family Furnishinidae. Fossils can be found in the Wheeler Shale Cambrian (c. 507 Mya) fossil locality in Utah, United States. Kambrische
Hertzina
Extinct genus of bacteria
modern cladophoran green algae. It is known from the Middle Cambrian Burgess Shale and Early Cambrian deposits from the Czech Republic. It comprises a dense
Marpolia
Organic-rich fine-grained sedimentary rock containing kerogen
Oil shale is an organic-rich fine-grained sedimentary rock containing kerogen (a solid mixture of organic chemical compounds) from which liquid hydrocarbons
Oil_shale
Extinct order of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Phacopida
Genus of trilobites (fossil)
tenuis (Illing, 1916) from the lower Tomagnostus fissus Biozone in the Abbey Shale Formation of Nuneaton, central England and the coeval lower Menevia Formation
Peronopsis
; Hu, J.; Zhang, J. (2026). "Late Ordovician ostracods from the "butter shale" Lagerstätten of the Cincinnati Arch region, Ohio, USA". Journal of Paleontology:
2026 in arthropod paleontology
2026_in_arthropod_paleontology
Extinct Cambrian family of animals
Burgess Shale and Spence Shale of western Laurentia. Skeemella has been found in the Pierson Cove and Marjum Formations, both above the Wheeler Shale and
Banffiidae
Extinct stem-arthropod species found in Cambrian fossil deposits
extinct stem group marine arthropod found in the Lagerstätte of the Burgess Shale (Middle Cambrian, 505 million years ago) of British Columbia. Opabinia was
Opabinia
Fifth stage of Cambrian and first stage of the Miaolingian
S. Lieberman (2023). "New soft-bodied panarthropods from diverse Spence Shale (Cambrian; Miaolingian; Wuliuan) depositional environments". Journal of
Wuliuan
Extinct genus of Cambrian Arthropod
community as of 2006. The genus is also known from Langston Formation, and Wheeler Shale of North America, as well as Chengjiang Biota of China (M. sinica).
Mollisonia
Geologic formation in Greenland
E. kingii fossil from the Wheeler Shale
Henson_Gletscher_Formation
Process for extracting oil from oil shale
Shale oil extraction is an industrial process for unconventional oil production. This process converts kerogen in oil shale into shale oil by pyrolysis
Shale_oil_extraction
Animals of the phylum Porifera
Cambrian Yuanshan Formation (Chengjiang, China) and the middle Cambrian Wheeler Shale (Utah, USA) and paleobiological implications". Journal of Paleontology
Sponge
Extinct group of artiopods
including but not limited to the Burgess Shale, Wheeler Shale, Chengjiang biota, Paseky Shale, Emu Bay Shale, Fezouata Formation, Llanfallteg Formation
Vicissicaudata
Extinct genus of arthropod
mythology. Multiple species had been recovered from the Burgess Shale Formation, Canada, Wheeler Shale and Marjum Formation, United States, and Valdemiedes Formation
Caryosyntrips
City in Utah, United States
relatively common in the region west of Delta (part of the House Range's Wheeler Shale). Several local companies maintain fossil dig areas where they allow
Delta,_Utah
Extinct family of Cambrian organisms
Cambrian Yuanshan Formation (Chengjiang, China) and the Middle Cambrian Wheeler Shale (Utah, USA) and paleobiological implications". Journal of Paleontology
Chancelloriidae
Extinct superfamily of trilobite
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Proetoidea
Group of extinct worm-like animals with legs
discovered in the Canadian Burgess Shale, and Hallucigenia, known from both the Chenjiang Maotianshan Shale and the Burgess Shale. Aysheaia pedunculata has a
Lobopodia
Extinct superfamily of trilobites
Ptychoparid trilobites of the Lower-Middle Cambrian boundary interval, Pioche Shale, southeastern Nevada, Journal of Paleontology, 604-630 Landing E., Johnson
Ptychoparioidea
Species of trilobite
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Phacops_fecundus
Family of extinct molluscs
Cambrian Yuanshan Formation (Chengjiang, China) and the Middle Cambrian Wheeler Shale (Utah, USA) and paleobiological implications". Journal of Paleontology
Halkieriid
Extinct order of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Ptychopariida
J. W. (1973). "Notobrachyops picketti, a brachyopid from the Ashfield Shale, Wiannamatta Group, New South Wales". Journal of Paleontology. 47 (6): 1094–1101
List_of_fossil_sites
Extinct genus of radiodonts
are known from Qingjiang biota in Hubei, China, Pioche Shale in Nevada, USA, and Wheeler Shale in Utah, USA. Huangshandongia yichangensis and Liantuoia
Hurdia
Second age of the Miaolingian epoch
Drum Mountains. The section is an outcrop of the Wheeler Formation, a succession of calcareous shales. The precise base of the Drumian is a laminated limestone
Drumian
Extinct order of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Harpetida
Shale known for exceptional preservation of mid-Cambrian organisms
The Burgess Shale of British Columbia is famous for its exceptional preservation of mid-Cambrian organisms. Around 69 other sites have been discovered
Burgess Shale-type preservation
Burgess_Shale-type_preservation
Genus of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Dinea
Extinct order of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Corynexochida
Extinct family of trilobites
interstrictus, until recently known as Peronopsis interstricta, from the Wheeler Shale, Utah Scientific classification Kingdom: Animalia Phylum: Arthropoda
Peronopsidae
7200°N 109.9647°E / 28.7200; 109.9647 Drumian ~504.5 Drumian section Wheeler Shale, Utah, U.S.A. Biologic: First appearance of Ptychagnostus atavus (Trilobite)
List of Global Boundary Stratotype Sections and Points
List_of_Global_Boundary_Stratotype_Sections_and_Points
Lead Bell Shale Cambrian Lehman Formation Ordovician Lodgepole Limestone Carboniferous Lone Mountain Formation Silurian Lower Wheeler Shale Cambrian Madison
List of fossiliferous stratigraphic units in Utah
List_of_fossiliferous_stratigraphic_units_in_Utah
found at Maotianshan Shales, whose most famous assemblage of organisms are referred to as the Chengjiang biota. The Maotianshan Shales are a series of Early
Paleobiota of the Maotianshan Shales
Paleobiota_of_the_Maotianshan_Shales
Extinct genus of trilobites
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Bumastus
Extinct species of trilobite
Silica Shale Formation Takaka terrane Walcott–Rust quarry Wenlock Group Wheeler Shale Wolchov River Wren's Nest Morphology Exoskeleton Tagmata Cephalon Thorax
Eldredgeops_rana
American paleontologist (1908–1992)
"Pseudoarctolepis sharpi, n. Gen., n. Sp. (Phyllocarida), from the Wheeler Shale (Middle Cambrian) of Utah". Journal of Paleontology. 30 (1): 9–14. JSTOR 1300370
Kenneth_Edward_Caster
depositional history of the Neoproterozoic Shaler Supergroup, Amundsen Basin, NW Canada: Rhenium-osmium dating of black shales from the Wynniatt and Boot Inlet
List_of_lagerstätten
Glacier in Nevada, United States
Wheeler Peak Glacier is a glacier situated at the base of Wheeler Peak within Great Basin National Park in the U.S. state of Nevada. It is the only glacier
Wheeler_Peak_Glacier
Proposed clade of extinct Lophotrochozoa
Cambrian Yuanshan Formation (Chengjiang, China) and the Middle Cambrian Wheeler Shale (Utah, USA) and paleobiological implications". Journal of Paleontology
Halwaxiida
Asaphiscus wheeleri, a trilobite from the Cambrian Wheeler Shale of Utah
List_of_trilobite_genera
WHEELER SHALE
WHEELER SHALE
Surname or Lastname
English
English : occupational name for a boatman or boatbuilder, from an agent derivative of Middle English kele ‘ship’, ‘barge’ (from Middle Dutch kiel).Americanized spelling of German Kühler, from a variant of an old personal name (see Keeling) or a variant of Kuhl.
Surname or Lastname
English
English : habitational name for someone from Weoley Castle in West Midlands (formerly in Worcestershire), named with Old English wēoh ‘(pre-Christian) temple’ + lēah ‘(woodland) clearing’, or from Weeley in Essex, which is named with Old English wilig ‘willow’ + lēah.
Surname or Lastname
English
English : variant of Wheatley.
Surname or Lastname
English
English : perhaps a variant spelling of Whit(t)la, itself a variant of Whitley.
Surname or Lastname
English
English : from Anglo-Norman French chivere, chevre ‘goat’ (Latin capra ‘nanny goat’), applied as a nickname for an unpredictable or temperamental person, or a metonymic occupational name for a goatherd.Born in London in about 1614, the son of spinner William Cheaver, Ezekiel Cheever came to Boston in June 1637. After a brief sojourn in New Haven, CT, he was master of the Boston Latin School from 1670 until his death in 1708. He had twelve children; his youngest son, also called Ezekiel, was the clerk to the court in the infamous Salem witchcraft trials of 1692.
Boy/Male
Christian & English(British/American/Australian)
A Wheeler
Surname or Lastname
English
English : variant spelling of Whistler.
Surname or Lastname
North German
North German : habitational name for someone from Heeten in the Netherlands near Deventer.English : unexplained; perhaps a variant of Hayter. Compare Heater.
Surname or Lastname
English
English : variant spelling of Shearer.Possibly an Americanized form of German Schürer, a southern variant of Scheurer.
Boy/Male
English American
Wheel maker.
Surname or Lastname
German
German : nickname from the small medieval coin known as the häller or heller because it was first minted (in 1208) at the Swabian town of (Schwäbisch) Hall. Compare Hall.Jewish (Ashkenazic) : habitational name for someone from Schwäbisch Hall.German : topographic name for someone living by a field named as ‘hell’ (see Helle 3).English : topographic name for someone living on a hill, from southeastern Middle English hell + the habitational suffix -er.Dutch : from a Germanic personal name composed of the elements hild ‘strife’ + hari, heri ‘army’.Jewish (Ashkenazic) : nickname for a person with fair hair or a light complexion, from an inflected form, used before a male personal name, of German hell ‘light’, ‘bright’, Yiddish hel.
Surname or Lastname
English
English : unexplained.
Boy/Male
Australian, British, English
A Driver; Wheel Maker
Surname or Lastname
English
English : origin uncertain; possibly an altered spelling of Wheelhouse, reflecting a local pronunciation.
Surname or Lastname
English
English : origin uncertain. Perhaps a variant of Wheeless, or of Wheels, from Old English hwēol ‘wheel’, and so a topographic name for someone who lived near a waterwheel, or a metonymic occupational name for someone in charge of one.
Surname or Lastname
English
English : variant spelling of Weller.
Surname or Lastname
English
English : variant of Whorley.
Surname or Lastname
English
English : probably a variant of Mellor. Compare Mealor, Mealer.
Surname or Lastname
English
English : occupational name for a maker of wheels (for vehicles or for use in spinning or various other manufacturing processes), from an agent derivative of Middle English whele ‘wheel’. The name is particularly common on the Isle of Wight; on the mainland it is concentrated in the neighboring region of central southern England.A founder of Salisbury, NH, in 1634 was John Wheeler.
Girl/Female
Tamil
Elethia | à®à®²à¯‡à®¤à®¿à®¯à®¾
Healer
WHEELER SHALE
WHEELER SHALE
WHEELER SHALE
WHEELER SHALE
WHEELER SHALE
WHEELER SHALE
WHEELER SHALE