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Genus of vascular plants in the family Isoetaceae
timing of the origin of modern Isoetes is subject to considerable uncertainty. The name of the genus may also be spelled Isoëtes. The diaeresis (two dots over
Isoetes
Circumpolar species of quillwort
Bulgaria. NatureServe. "Isoetes lacustris". NatureServe Explorer. Arlington, Virginia. Retrieved 2007-12-19. "Synonymy - Isoëtes lacustris". Northern Ontario
Isoetes_lacustris
Western Asian species of quillwort
T164351A5837032.en. Retrieved 13 April 2022. "Isoetes olympica A.Braun". www.gbif.org. Retrieved 2022-03-05. "Isoetes olympica A.Br. | Plants of the World Online
Isoetes_olympica
Western Australian species of quillwort
Isoetes australis is quillwort (in the family, Isoetaceae) and was first described in 1943 by Samuel Williams. It is endemic to south-west Western Australia
Isoetes_australis
South African endemic quillwort species
Isoetes stellenbossiensis, the Stellenbosch quillwort or Cape Flats quillwort, is a species of plant from South Africa. This tufted geophyte has three-pointed
Isoetes_stellenbossiensis
Ecuadorian endemic species of quillwort
Isoetes ecuadoriensis is a species of plant in the family Isoetaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical high-altitude
Isoetes_ecuadoriensis
Western North American species of quillwort
Isoetes nuttallii, or Nuttall's quillwort, is a species of quillwort, a type of lycopod. It is native to shallow waters and other wet habitats of western
Isoetes_nuttallii
North American species of quillwort
originally believed to be a member of Isoetes hieroglyphica because of their similar megaspore structure. NRCS. "Isoetes acadiensis". PLANTS Database. United
Isoetes_acadiensis
Eastern Asian species of quillwort
Isoetes sinensis, the narrow quillwort, is a species of plant in the family Isoetaceae. Isoetes sinensis is native to China and Korea. Its natural habitat
Isoetes_sinensis
Southeastern United States species of quillwort
it as a synonym of Isoetes valida, but others treat it as a full species. Hassler, Michael & Schmitt, Bernd (November 2019), "Isoetes caroliniana", Checklist
Isoetes_caroliniana
Southeastern US species of quillwort
Isoetes melanospora, commonly known as black-spored quillwort or black-spored Merlin's grass, is a rare and endangered aquatic lycophyte endemic to the
Isoetes_melanospora
Species of spore-bearing plant
Isoetes × eatonii, or Eaton's quillwort, is a hybrid between I. engelmannii and I. tenella. It can be found in ponds and slow moving rivers in Canada
Isoetes_×_eatonii
European and African species of quillwort
Isoetes histrix, the land quillwort, is an aquatic pteridophyte native to the Mediterranean region, northwestern Africa, and the coasts of western Europe
Isoetes_histrix
Species of spore-bearing plant
"Isoetes tegetiformans". Flora of North America. Vol. 2. New York & Oxford: Oxford University Press. Wikimedia Commons has media related to Isoetes tegetiformans
Isoetes_tegetiformans
Extinct genus of vascular plants
impression known as the "ligular pit" for its similarities to the ligule of Isoetes. In some leaf-cushions a second depression was present above the ligular
Lepidodendron
South African quillwort species
Isoetes capensis, the cape quillwort, is a species of quillwort from South Africa. The cape quillwort is a tufted geophyte with horny toothed scales.
Isoetes_capensis
Western North American species of quillwort
Isoetes occidentalis, the western quillwort, is a species of quillwort in the family Isoetaceae. It can be found in aquatic habitats of coastal Alaska
Isoetes_occidentalis
Temperate Northern Hemisphere species of quillwort
plant lack the stomata present in North American populations. Isoetes muricata and Isoetes echinospora var. braunii refer to the North American plants,
Isoetes_echinospora
New Zealand endemic species of quillwort
fine sediments or coarse sand.[citation needed] Isoetes alpina Kirk, nzflora, retrieved 30 May 2019 Isoetes alpina, New Zealand Plant Conservation Network
Isoetes_alpina
Temperate North American species of quillwort
New Jersey, and Tennessee. "Isoetes melanopoda Gay & Durieu". www.worldfloraonline.org. Retrieved 2021-01-16. "Isoetes melanopoda J.Gay & Durieu". Plants
Isoetes_melanopoda
Eastern North American species of quillwort
2020. "Isoetes riparia Engelm. ex A. Braun". World Flora Online. The World Flora Online Consortium. n.d. Retrieved September 26, 2020. NRCS. "Isoetes ripara"
Isoetes_riparia
South African endemic quillwort species
Stephansen, for whom it was named. It was reduced to a variety of Isoetes capensis as Isoetes capensis var. stephanseniae in 1985, but more recent treatments
Isoetes_stephanseniae
Eastern North American species of quillwort
of Isoetes butleri die back at the end of spring, when its habitat becomes hot and dry. It is then reduced to a below ground corm. NRCS. "Isoetes butleri"
Isoetes_butleri
Species of plant
Isoetes araucaniana is a species of aquatic lycophyte in the family Isoetaceae. It is endemic to Chile, distributed between the Biobio and Araucania regions
Isoetes_araucaniana
Arts. Retrieved 2026-07-14. "Isoetes Japonica Live Aquatic Plant". Glass Aqua. Retrieved 2026-07-14. "Quillwort (Isoetes lacustris)". Aqua Imports. Retrieved
List of freshwater aquarium plant species
List_of_freshwater_aquarium_plant_species
Species of spore-bearing plant
"Isoetes macrospora Durieu". Plants of the World Online. The Trustees of the Royal Botanic Gardens, Kew. n.d. Retrieved July 23, 2025. NRCS. "Isoetes macrospora"
Isoetes_macrospora
South African species of quillwort
Isoetes toximontana, the Gifberg quillwort, is a plant species native to the Northern Cape and Western Cape regions of South Africa. It is known from
Isoetes_toximontana
Species of spore-bearing plant
introduction. Some sources consider Isoetes caroliniana a synonym of this species, others regard it as a separate species. "Isoetes valida". NatureServe Explorer
Isoetes_valida
Eastern North American species of quillwort
Isoetes engelmannii is a species of aquatic plant in the family Isoetaceae. It is referred to by the common names Engelmann's quillwort or Appalachian
Isoetes_engelmannii
Mississippian endemic species of quillwort
panicgrass. Isoetes mississippiensis only occurs in two known locations, neither of which is in a protected area. The rootstock of Isoetes mississippiensis
Isoetes_mississippiensis
Eastern North American species of quillwort
Isoetes appalachiana, commonly known as the Appalachian quillwort (not to be confused with its close relative Isoetes engelmannii, which shares the same
Isoetes_appalachiana
Species of plant in the family Isoetaceae
"Isoetes tuckermanii A. Braun ex Engelm". World Flora Online. The World Flora Online Consortium. n.d. Retrieved September 26, 2020. NRCS. "Isoetes tuckermanii"
Isoetes_tuckermanii
Southeastern US species of quillwort
Isoetes louisianensis, the Louisiana quillwort, is a small, grass-like aquatic plant of the family Isoetaceae. It is "one of the rarest quillworts in
Isoetes_louisianensis
Extinct genus of spore-bearing plants
weed in lake and pond sedimentary environments, like living Isoetes. Unlike living Isoetes, Tomiostrobus formed closed cones with sporophylls that were
Tomiostrobus
South African endemic species of quillwort
Isoetes eludens is an aquatic plant in the genus commonly known as quillwort that is native to the Kamiesberg Mountains in Namaqualand, South Africa.
Isoetes_eludens
Western Australian species of quillwort
Isoetes brevicula is a quillwort species native to Western Australia. It is a cormous, perennial, herb or (fern ally), to 1 cm (0.39 in) high, stock 3-lobed;
Isoetes_brevicula
Taiwanese endemic species of quillwort
Isoetes taiwanensis is a species of plant in the family Isoetaceae. It is endemic to Taiwan, and the only species of quillwort there. Isoetes taiwanensis
Isoetes_taiwanensis
Southern European species of quillwort
April 2022. "Isoetes heldreichii Wettst. | Plants of the World Online | Kew Science". Plants of the World Online. Retrieved 27 May 2021. "Isoetes heldreichii"
Isoetes_heldreichii
Species of spore-bearing plant
of the genus Isoetes, which contains living quillworts, though it differs from living Isoetes in some aspects, with modern forms of Isoetes possibly emerging
Isoetes_beestonii
Order of free-sporing vascular plants
more closely related to "aborescent lycophytes" that to Isoetes. Fossilised specimens of Isoetes beestonii have been found in rocks dating to the latest
Isoetales
Italian endemic species of quillwort
Isoetes malinverniana, the Piedmont quillwort, is a species of quillwort. It is critically endangered. It is found in Italy. There is a breeding program
Isoetes_malinverniana
French screenwriter and writer (1945–2026)
(photography) (Isoète, 1991) Cherbourg, with Natacha Hochman (photography) (Isoète, 1992) Presqu'île de lumière, with Patrick Courault (photography) (Isoète, 1996)
Didier_Decoin
Index of plants with the same common name
name for several plants and may refer to: Isoetes appalachiana, native to the eastern United States Isoetes engelmannii, native to eastern North America
Appalachian_quillwort
Family of spore-bearing plants
Isoetaceae is a family including living quillworts (Isoetes) and comparable extinct herbaceous lycopsids (Tomiostrobus). It was previously thought to
Isoetaceae
North American species of quillwort
Isoetes howellii, or Howell's quillwort, is a species of quillwort, a type of lycophyte. It is an aquatic plant native to North America. Isotes howelii
Isoetes_howellii
Class of vascular plants
(Lycopodiales) are homosporous, but the genera Selaginella (spikemosses) and Isoetes (quillworts) are heterosporous, with female spores larger than the male
Lycopodiopsida
North American species of quillwort
Isoetes bolanderi, or Bolander's quillwort, is a species of quillwort, a type of lycophyte. This aquatic plant is native to high altitude regions of the
Isoetes_bolanderi
Species of plant native to Australia
Arten der Gattung Isoetes". Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin 1868. p. 541. "Isoetes muelleri A.Braun".
Isoetes_muelleri
Species of aquatic plant
to Isoetes. The leaves of Lobelia dortmanna are, however, easily distinguishable from those of other isoetids in having only two air-canals (Isoetes having
Lobelia_dortmanna
Lake in Maine, United States
was formerly known as Isoetes macróspora var. heteróspora (or Isoetes heterospora) but is now considered to be a synonym for Isoetes lacustris, which is
Jordan_Pond
Species of moth
aquatic and have been recorded feeding on Lobelia dortmanna, Isoetes tuckermani and Isoetes muricata. They have a yellow head. Full-grown larvae reach a
Eoparargyractis_plevie
Extinct genus of lycopsid plants in the Order Isoetales
Pleuromeia is an extinct genus of lycophytes related to modern quillworts (Isoetes). Pleuromeia dominated vegetation during the Early Triassic across Eurasia
Pleuromeia
Extinct genus of spore-bearing plants
stigmarian rootlets branched dichotomously from the rhizomorphs similar to Isoetes, and spread throughout the coal swamp forest areas where the lycopods were
Sigillaria
Elaphoglossum gracilipes Elaphoglossum polytrichum Isoetes heldreichii, Pindus quillwort Isoetes sinensis Melpomene brevipes Extinct in the wild species
List of recently extinct plants
List_of_recently_extinct_plants
English name Scientific name Status Spring quillwort Isoetes echinospora Native Land quillwort Isoetes histrix Native Quillwort Isoetes lacustris Native
List of ferns and fern allies of Great Britain and Ireland
List_of_ferns_and_fern_allies_of_Great_Britain_and_Ireland
Genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Diphasium
blue-eyed-grass Isoetes acadiensis — Acadian quillwort Isoetes bolanderi — Bolander's quillwort Isoetes echinospora — spinyspore quillwort Isoetes engelmannii
List of Canadian plants by family I–K
List_of_Canadian_plants_by_family_I–K
Mountain and park in Georgia, United States
listed rare and endangered plant species, such as black-spored quillwort (Isoetes melanospora) and pool sprite (also called snorkelwort, Gratiola amphiantha)
Stone_Mountain
Species of quillwort
Isoetes maritima, the maritime quillwort, is a quillwort in the Isoetaceae family. It is native to Alaska, British Columbia, and Washington state. It
Isoetes_maritima
Metabolic process
genera, including: Isoetes, Crassula, Littorella, Sagittaria, and possibly Vallisneria, being found in a variety of species e.g. Isoetes howellii, Crassula
Crassulacean_acid_metabolism
Extinct genus of lycopsid plants
for being a potential bridge between the Triassic Pleuromeia and extant Isoetes. It is known from one collection of nearly 270 specimens that include many
Nathorstiana
Type of growth in plants
living pteridophytes this feature is extremely rare, only occurring in Isoetes. In many vascular plants, secondary growth is the result of the activity
Secondary_growth
Production of spores of two different sizes and sexes by several groups of land plants
differentiation. Four extant groups of plants are heterosporous; Selaginella, Isoetes, Salviniales and seed plants. Heterospory evolved due to natural selection
Heterospory
Extinct genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Baragwanathia
Lake in New Brunswick, Canada
stagnalis is found mainly along the lake's margin, composed chiefly of Isoetes echinospora; Sagittaria montevidensis; Potamogeton gramineus; Potamogeton
Chamcook_Lake
Extinct order of vascular tree-like plants
mosses", they are believed to be more closely related to extant quillworts (Isoetes) based on xylem (and consequently sometimes placed within the Isoetales
Lepidodendrales
Group of vascular plants
Isoëtes lacustris, a quillwort, from the Isoetopsida
Fern_ally
Index of plants with the same common name
Merlin's grass is a common name for several plants and may refer to: Isoetes lacustris Isoetes tegetiformans, endemic to the U.S. state of Georgia This page
Merlin's_grass
Extinct genus of lycopsid plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Chaloneria
UK plant community type
Ranunculetum baudotii Br.-Bl. 1952 A22 Littorella uniflora - Lobelia dortmanna community A23 Isoetes lacustris/setacea community A24 Juncus bulbosus community.
Aquatic communities in the British National Vegetation Classification system
Aquatic_communities_in_the_British_National_Vegetation_Classification_system
Second period of the Mesozoic Era
known species to represent all major morphological features of modern Isoetes. More primitive forms such as Nathorstiana, which retain an elongated stem
Jurassic
Isoetes engelmannii Isoetes giessii Isoetes howellii Isoetes lacustris, common quillwort Isoetes lechleri Isoetes maritima Isoetes nuttallii Isoetes palmeri
List_of_least_concern_plants
Spends part of their time in water, or grows partially submerged in water
semiaquatic horsetail, Equisetum fluviatile Semiaquatic quillworts, such as Isoetes melanospora Semiaquatic club mosses, such as Lycopodiella inundata Semiaquatic
Semiaquatic
Small inland lake in Scotland
Flora include Littorella uniflora, Lobelia dortmanna, Juncus bulbosus and Isoetes. Brown trout and Arctic charr are to be found in the loch. Dippers flip
Loch_Einich
Genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Spinulum
Class of vascular plants
; et al. (2010). "Complete plastome sequences of Equisetum arvense and Isoetes flaccida: implications for phylogeny and plastid genome evolution of early
Fern
fadeniana Isoetes dispora Isoetes heldreichii, Pindus quillwort (possibly extinct) Isoetes hewitsonii Isoetes malinverniana, Piedmont quillwort Isoetes nigroreticulata
List of critically endangered plants
List_of_critically_endangered_plants
of rootlets. A similar construction is observed in the extant lycopod Isoetes, and this appears to be evidence that roots evolved independently at least
Evolutionary history of plants
Evolutionary_history_of_plants
Nature reserve
the botanist Edith Layard Stephens. She intended it to preserve the rare Isoetes capensis plant, which exists nowhere else on Earth. After subsequent additions
Edith_Stephens_Wetland_Park
Woodsia fragilis (Trev.) Moore - (rare) Isoetes lacustris L. - (rare) Isoetes maritima Underw. - (declining) Isoetes setacea Durieu - (declining) Juniperus
List of the vascular plants in the Red Data Book of Russia
List_of_the_vascular_plants_in_the_Red_Data_Book_of_Russia
Lake in Schleswig-Holstein, Germany
nutrient-poor waters, including water lobelia (Lobelia dortmanna), brazengrass (Isoetes lacustris), shoreweed (Littorella uniflora), and alternate water-milfoil
Bültsee
Genus of vascular plants in the family Lycopodiaceae
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Lycopodium
Genus of vascular plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Huperzia
German botanist (1805-1877)
morphology). 1862: Zwei deutsche Isoetesarten (Two German Isoëtes species). 1863: Über Isoetes (On quillworts). 1865: Beitrag zur Kenntnis der Gattung Selaginella
Alexander_Braun
float on the surface of water, with members of the group found worldwide. Isoetes is a cosmopolitan genus of lycophyte known as the quillworts. Juncus is
List_of_wetland_plants
Genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Phylloglossum
Ranunculus kazusensis (매화마름): It lives in the southern province and west coast. Isoetes (물부추): It grows in shallow water. It was found in Pyeongtaek in 1942; it
Wildlife_of_South_Korea
Broadly circumscribed group of spore bearing plants
Wisconsin. Lycopodium dendroideum, a modern member of the Lycopodiales Isoetes melanospora, a modern member of the Isoetales Restoration of Pleuromeia
Lycophyte
Extinct genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Asteroxylon
Topics referred to by the same term
Quillwork, traditional Native American art form Quillwort, common name of Isoetes This disambiguation page lists articles associated with the title Quill
Quill_(disambiguation)
wetland plants named for their superficial similarity to the quillworts, Isoetes. They occur in wetlands and on shorelines with low nutrient availability
Isoetid
Protected area in northern Alberta, Canada
larger trees around the lakes. The rare, spore-bearing plant quillwort (Isoetes echinospora) was discovered in the area in 2004. The park is within the
Birch Mountains Wildland Provincial Park
Birch_Mountains_Wildland_Provincial_Park
American botanist (1889-1989)
(1889–1989) was an American botanist who specialized in the study of lilies and Isoetes. She discovered and described the Chicago-endemic flowering plant species
Norma_Etta_Pfeiffer
Vernicia fordii. There are 112 national first-level protected plants: Isoetes sinensis, Davidia involucrata, Cycad, Ginkgo, Cathaya, Metasequoia glyptostroboides
Geography_of_Hunan
Genus of spore-bearing plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Dendrolycopodium
Genus of plants
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Palhinhaea
Protected area in Estonia
plants growing in the area are Pulsatilla patens, Sparganium angustifolium, Isoetes echinospora and Oxytropis sordida (the only place in Estonia where the
Põhja-Kõrvemaa_Nature_Reserve
Elaphoglossum serpens (no common name) Isoetes louisianensis (Louisiana quillwort) Isoetes melanospora (black spored quillwort) Isoetes tegetiformans (mat-forming
List of endangered plants of North America
List_of_endangered_plants_of_North_America
genus Isoetes in North America. In: Transactions of the Academy of Science of St. Louis. Vol 4, 1882, S. 358–390, (online). The genus Isoetes in North
George_Engelmann_bibliography
Genus of vascular plants in the clubmoss family Lycopodiaceae
†Takhtajanodoxa †Nathorstiana †Lepacyclotes †Isoetites †Skilliostrobus †Tomiostrobus Isoetes †Lepidodendrales Lepidodendron Lepidophloios Ulodendron Sublepidophloios
Diphasiastrum
ISOETES
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ISOETES
n.
One of the exceedingly minute spores found in certain flowerless plants, as Selaginella and Isoetes, which bear two kinds of spores, one very much smaller than the other. Cf. Macrospore.
n.
Any plant or species of the genus Isoetes, cryptogamous plants with a cluster of elongated four-tubed rushlike leaves, rising from a corm, and containing spores in their enlarged and excavated bases. There are about seventeen American species, usually growing in the mud under still, shallow water. So called from the shape of the shape of the leaves.
n.
A thin membrane surrounding the sporocarps of quillworts Isoetes).